Notophthalmus viridescens (inset photo courtesy of and copyright by Will Brown)

INTRODUCTION


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Red Eft (from University of Illinoise at Urbana-Champaign Herp Picture website: http://gto.ncsa.uiuc.edu/pingleto/herps/sally.html)

The Eastern Newt, Notophthalmus viridescens (Rafinesque), has a wide range throughout the eastern half of North America. It is one of the most commonly found species of salamanders in Connecticut. Throughout most of its range, this species has an interesting life cyle, usually comprised of three distinct stages -- an aquatic larval stage, a terrestrial juvenile stage, and aquatic adult stage. The diurnal habit and its brilliant color make the terrestrial eft a familiar sight in moist woodlands in many parts of the species' range.

Please use the topic bar on the left to choose the subject that interests you. For image description and credit, please place the pointer over the image for the pop-up information box. For more information on the Eastern Newt, please refer to Reference and Online Resources pages for related literatures and internet links. Interested in seeing a specimen? Check the Museums page for a list of natural history museums with specimens of Notophthalmus viridescens in their herpetology collection.


SUBSPECIES

Four subspecies of Notophthalmus viridescens (Rafinesque) have been recognized on the basis of dorsal color patterns:

  • Notophthalmus viridescens viridescens (Rafinesque)
    Red-spotted Newt, Spotted Newt, Common Newt, Red Eft (terrestrial juvenile stage)
    Type-locality: Lake George and Lake Champlain, New York
  • Notophthalmus viridescens dorsalis (Harlan)
    Broken-striped Newt
    Type-locality: Vicinity of Camden, South Carolina
  • Notophthalmus viridescens louisianensis (Wolterstorff)
    Central Newt, Louisiana Newt
    Type-locality: New Orleans, Louisiana
  • Notophthalmus viridescens piaropicola (Schwartz & Duellman)
    Peninsula Newt
    Type-locality: Collier County, Florida
The Red-spotted Newt, N. v. viridescens, is the only subspecies found in Connecticut. Click here for the distribution of subspecies of Notophthalmus viridescens.


CLASSIFICATION

Notophthalmus viridescens is classified as follows:

  • KINGDOM Animalia
  • PHYLUM Chordata
  • CLASS Amphibia
  • SUBCLASS Lissamphibia
  • ORDER Caudata
  • FAMILY Salamandridae
The family Salamandridae includes 53 extant species placed in 15 genera distributed primary in Europe and Asia; two genera are endemic to North America. Taricha encompasses three species restricted to the Pacific coast region, while the three species of Notophthalmus occur in the eastern half of United States and Canada, of which N. viridescens is the sole representive in the New England area. Based on fossil and biochemical data, the two genera are estimated to have diverged at least 20 million years ago (Reilly 1990).

Salamandrids (newts) lack costal grooves and slimy skin that characterize most other salamanders. All salamandrids have well-developed limbs, and many aquatic species have dorsal and caudal fins. Members of Salamandridae all have toxic skin secretions that serve as defense against predators. The larvae have four pairs of gill slits and large external gills (Duellman & Trueb 1994).


TAXONOMIC HISTORY

The following summary of synonymies and references for Notophthalmus viridescens follows that given by Mecham (1967):

  • Notophthalmus viridescens (Rafinesque)
  • Triturus (Diemictylus) viridescens Rafinesque, 1820:5. Type locality, "in Lake George, Lake Champlain, the springs and brooks of the neighborhood." No type designated and collector not given. Diemictylus proposed as a subgenus.
  • Triturus (Notophthalmus) miniatus Rafinesque, 1820:5. Type-locality, "states of New York, New Jersey, Connecticut, Vermont, etc."; restricted to "borders of Lake Champlain" by Schmidt, 1953:24. Type not designated and collector not given. Description based on eft stage. Notophthalmus proposed as a subgenus.
  • Notopthalmus viridescens: Baird, 1850:284.
  • Diemyctylus viridescens: Hallowell, 1856:11.
  • Notophthalma viridescens: Gray, 1858:138.
  • Triton viridescens: Strauch, 1870:50.
  • Molge viridescens: Boulenger, 1882:21.
  • Notophthalmus viridescens: Gill, 1907:256.
  • Diemictylus viridescens: Gill, 1907:256. Given as a synonym of Notophthalmus viridescens.

Eft on the forest floor (photo courtesy of Joyce Gross, from UC Berkeley AmphibiaWeb) Rafinesque is credited as the first author to name the Eastern Newt in 1820, where he described the adult and eft stages as separate species belonging to the genus Triturus : Triturus (Diemictylus) viridescens (1820:5, line 8) and Triturus (Notophthalmus) miniatus (1820:5, lines 26-27). However, his name appears in parenthesis since the names he designated are not the currently accepted scientific name for the species. His two proposed subgenera were nonetheless the earliest names applicable to the Eastern Newts and both admissible as the generic name when subsequent authors (preceded by colons) revised to form a new genus based on the union of the two subgenera, which in fact represented the same species. By the International Rules on Zoological Nomenclaure,

"A genus formed by the union of two or more genera or subgenera takes the oldest valid generic or subgeneric name of its components. If the names are of the same date, that selected by the first reviser shall stand."

Thus, selection of the "first reviser" takes precedence over line priority (i.e. Diemictylus first mentioned by Rafinesque in line 8 while Notophthalmus first appeared in lines 25-26 on the same page of the same 1820 published account). The first reviser happened to be Baird (1850:284), who selected Notophthalmus (with an invalid spelling change to Notopthalmus) and synonymized with it the name Diemictylus, six years before Hallowell (1856:11) chose Diemictylus (with an invalid spelling change to Diemyctylus). Baird's selection was upheld by Gray (1858:138) who erraneously spelled it as Notophthalma. The specific name viridescens was given a separate consideration from the generic name and determined to be a senior synonym of miniatus on grounds both of line priority and of first reviser. For a summary of the nomenclatural history of the Eastern Newt, see Smith (1953).


ETYMOLOGY

Green dorsum of adult (from Nova Scotia Museum of Natural History Salamanders website: http://www.ednet.ns.ca/educ/museum/mnh/nature/salamand/rspot.htm) The specific name viridescens (Latin, viridis, "green") refers to the greenish color often found in the adults of this species. The supspecific name dorsalis (directly from Latin, "pertaining to the back") presumably alludes to the broken red lines on the dorsum of the broken-striped newt. The type locality for the central newt is in the state of Louisiana, hence the subspecific name louisianensis. The peninsula newt is often found inhabiting beds of water hyacinths (Piaropus), yielding the name piaropicola (Mecham 1967).

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